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When the cell of a typical apple is ruptured – for example, by biting, slicing or bruising – polyphenol oxidase (PPO) found in one part of the cell mixes with polyphenolics found in another part of the cell. Polyphenolics are one of the many types of chemical substrates that serve various purposes, including supplying its aroma and flavor.) When PPO and polyphenolics mix, brown-toned melanin is left behind.

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